104
GREAT DIVING BEETLES
DYTISCIDAE: DYTISCINAE
DYTISCIDAE: DYTISCINAE
subfamily
Dytiscinae
known species
380–400
distribution
Worldwide except Antarctica. Large species
are common in northern climates as well as
in the tropics
habitat
Fresh water, including ponds, lakes, rivers,
and some brackish areas near the coast. Can
be found in very small, recently formed pools
size
10–45 mm
diet
Carnivorous. Adults and larvae feed
on aquatic arthropods, worms, fish,
and amphibians. Can be a minor pest
in fish or shrimp farms
lthough comprising only 10 percent of the
species of diving beetles, the subfamily
Dytiscinae, known as the great diving beetles,
includes some of the largest and most conspicuous
of all aquatic beetles. The biggest species, at almost
2 in (5 cm), is Megadytes ducalis; until recently it was
known from only a single specimen apparently
found in a water-filled canoe along the Amazon
River in the late 1800s, and kept at the Natural
History Museum in London.
All Dytiscinae are rapacious hunters as adults
and larvae. The large, yellowish-brown, striped
larvae, called “water tigers,” have hollow, sickle-
shaped mandibles that are used to liquify their prey
with enzymes before sucking up its juices, and they
often attack larger animals, even fish. On reaching
full size, larvae crawl out of the water and pupate in
a hole in the ground, the fresh adult either returning
to the pond or dispersing in search of new water
bodies. Dispersing adults fly powerfully at night, and
may be attracted to lights, or drop down onto shining
objects such as cars or glasshouses, mistaking the
reflected moonlight for a water surface. They have
even been found at oases deep in the Sahara Desert.
left | Dytiscus marginalis The voracious larvae
of this European great diving beetle often eat
vertebrates such as fish, newts, and tadpoles.
A